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Sophia Myles (born 18 March 1980) played Jeanne-Antoinette Poisson in the Doctor Who story The Girl in the Fireplace. Prior to her appearance on Doctor Who, she was best known for appearing in costume dramas: she appeared in Underworld and as Lady Penelope in the remake of Thunderbirds, along with Ron Cook as Penelope's Driver, Parker. Myles also starred as Isolde in Tristan + Isolde and as Kate Breck in Hallam Foe.
Before she starred in Doctor Who, she was romantically involved with her The Life and Adventures of Nicholas Nickleby co-star Charles Dance, thirty-four years her senior.
During filming of The Girl in the Fireplace, she began dating David Tennant and the relationship continued for two years.[source needed] During this period, she assisted the Doctor Who production crew by reading the role of Donna Noble at the cast run-through for The Runaway Bride (Catherine Tate was not available). She also helped Tennant record some of his video diaries for the DVD box sets, appeared in a couple of them and also handled the camera.
The relationship between Myles and Tennant reportedly ended[source needed] around the time she relocated to America to co-star in the 2007-08 series Moonlight, which was cancelled after one season. Also around this time she appeared in the film Art School Confidential.
Like Tennant, she also appeared in an episode of Extras.