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The Red Dalek, also known as Supreme One and Dalek Supreme, was in command of the New Dalek Empire in 2009. It was also a variant of the Dalek Supreme model.
Biography
It commanded the invasion of Earth during the War in the Medusa Cascade and reported its victory to Davros though he was only interested if The Doctor had arrived. The Supreme One stated that there was no sign of Time Lord activity and that they were beyond the Doctors reach which fascinated Davros as it sounded as a declaration of victory which was impossible as the Daleks were elevated above emotion. (DW: The Stolen Earth)
Once its forces detected the Doctor's TARDIS, the Supreme One ordered it to be encased in a time prison which allowed it to be captured as it was depowered. The craft was transported to the Crucible where the Supreme One demanded that the Doctor and his companions emerge from their craft. When the door to the TARDIS closed trapping Donna Noble, the Doctor told the Supreme One to release her but the red Dalek stated it was not his action and that it was likely Time Lord trickery.
It commanded its forces to dump the TARDIS into the core of the Crucible in order to destroy it. As it disintegrated, the Supreme One asked the Doctor if he had felt anger, that if emotions made one stronger. Captain Jack Harkness then attacked the Supreme One with a pistol which did not harm the Dalek leader. The Supreme Dalek retaliated by using its energy weapon in an attempt to kill Jack, unaware that he was immortal. After which, it ordered the Doctor and Rose Tyler to be taken to the Vault.
The Doctor discovered in his meeting with Davros that the creator of the Daleks was not the master of the New Dalek Empire but it was in fact the Supreme One who was in charge. After being created, the Daleks turned on their creator (again) who made an arrangement with them where he would ensure their domination of the universe. This was to be done through the use of a Reality bomb which would be powered by the 27 planetary engine configuration of the captured worlds in the Medusa Cascade in conjunction with the Crucible's energy source. This would bring about the destruction of all existence except for the Daleks leaving them to be the masters of the universe. To ensure Davros's cooperation, the Supreme One locked Davros - along with the insane Dalek Caan - in the Vault of the Crucible.
The Supreme One then received the transmission from Martha Jones who threatened to destroy Earth with the Osterhagen key thus preventing the Daleks from completing their goal. The Daleks then used a Transmat to capture the Human. However, the arrival of Donna Noble and a second Doctor brought ruin to their plan as the reality bomb did not activate. A furious Supreme One entered the Vault where he angrily stated that Davros betrayed the Daleks to which an innocent Davros protested that it was Dalek Caan. The Supreme One ignored him and declared that all the Vault's occupants would be exterminated. He was only able to get a single shot off, however, before being hit by a blast from Captain Jack's energy weapon. The Supreme One screamed as it burst into flames and exploded. (DW: Journey's End)
Overview
The Red Dalek had red casing with golden globes and extra plating. It had a slightly deeper voice than the other Daleks and ranked even above Davros. As well as extra armour, the Red Dalek had a third indicator light which - as with its other lights - was mounted on a small plate.
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The Red Dalek had a clear dislike for Dalek Caan, calling him "the Abomination".